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FAMILY 64.–TODIDÆ. (1 Genus, 5 Species.)

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The Todies are delicate, bright-coloured, insectivorous birds, of small size, and allied to the Motmots, although externally more resembling flycatchers. They are wholly confined to the greater Antilles, the islands of Cuba, Hayti, Jamaica, and Porto Rico having each a peculiar species of Todus, while another species, said to be from Jamaica, has been recently described (Plate XVI. Vol. II. p. 67).

FAMILY 65.—MOMOTIDÆ. (6 Genera, 17 Species.)

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The Motmots range from Mexico to Paraguay and to the west coast of Ecuador, but seem to have their head-quarters in Central America, five of the genera and eleven species occurring from Panama northwards, two of the genera not occurring in South America. The genera are as follows :

Momotus (10 sp.), Mexico to Brazil and Bolivia, one species extending to Tobago, and one to Western Ecuador; Urospatha (1 sp.), Costa Rica to the Amazon ; Baryphthengus (1 sp.), Brazil and Paraguay ; Hylomanes (2 sp.), Guatemala ; Prionirhynchus (2 sp.), Guatemala to Upper Amazon; Eumomota (1 sp.), Honduras to Chiriqui.

FAMILY 66.—TROGONIDÆ. (7 Genera, 44 Species.)

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The Trogons form a well-marked family of insectivorous forest-haunting birds, whose dense yet puffy plumage exhibits the most exquisite tints of pink, crimson, orange, brown, or metallic green, often relieved by delicate bands of pure white. In one Guatemalan species the tail coverts are enormously lengthened into waving plumes of rich metallic green, as graceful and marvellous as those of the Paradise-birds. Trogons are tolerably abundant in the Neotropical and Oriental regions, and are represented in Africa by a single species of a peculiar genus. The genera now generally admitted are the following:

Trogon (24 sp.), Paraguay to Mexico, and west of the Andes in Ecuador; Temnotrogon (1 sp.), Hayti; Prionoteles (1 sp.), . Cuba (Plate XVII. Vol. II. p. 67); Apaloderma (2 sp.), Tropical and South Africa; Harpactes (10 sp.), the Oriental region, excluding China ; Pharomacrus (5 sp.), Amazonia to Guatemala ; Euptilotis (1 sp.), Mexico.

Remains of Trogon have been found in the Miocene deposits of France; and we are thus able to understand the existing distribution of the family. At that exceptionally mild period in the northern hemisphere, these birds may have ranged over all Europe and North America ; but, as the climate became more severe they gradually became restricted to the tropical regions, where alone a sufficiency of fruit and insect-food is found all the year round.

FAMILY 67.—ALCEDINIDÆ. (19 Genera, 125 Species.)

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The Kingfishers are distributed universally, but very unequally, over the globe, and in this respect present some of the most curious anomalies to be found among birds. They have their metropolis in the eastern half of the Malay Archipelago (our first Australian sub-region), from Celebes to New Guinea, in which district no less than 13 out of the 19 genera occur, 8 of them being peculiar; and it is probable that in no other equally varied group of universal distribution, is so large a proportion of the generic forms confined to so limited a district. From this centre kingfishers decrease rapidly in every direction. In Australia itself there are only 4 genera with 13 species ; the whole Oriental region has only 6 genera, 1 being peculiar; the Ethiopian also 6 genera, but 3 peculiar; and each of these have less than half the number of species possessed by the Australian region. The Palæarctic region possesses only 3 genera, all derived from the Oriental region; but the most extraordinary deficiency is shown by the usually rich Neotropical region, which possesses but a single genus, common to the larger part of the Eastern Hemisphere, and the same genus is alone found in the Nearctic region, the only difference being that the former possesses eight, while the latter has but a single species. These facts almost inevitably lead to the conclusion that America long existed without kingfishers; and that in comparatively recent times (perhaps during the Miocene or Pliocene period), a species of the Old World genus, Ceryle, found its way into North America, and spreading rapidly southward along the great river-valleys has become differentiated in South America into the few closely allied forms that alone inhabit that vast country-the richest in the world in

fresh-water fish, and apparently the best fitted to sustain a varied and numerous body of kingfishers.

The names of the genera, with their distribution and the number of species in each, as given by Mr. Sharpe in his excellent monograph of the family, is as follows:

Alcedo (9 sp.), Palæarctic, Ethiopian, and Oriental regions (but absent froin Madagascar), and extending into the Austro-Malayan sub-region ; Corythornis (3 sp.), the whole Ethiopian region ; Alcyone (7 sp.), Australia and the Austro-Malayan sub-region, with one species in the Philippine Islands ; Ceryle (13 sp.), absent only from Australia, the northern half of the Palæarctic region, and Madagascar ; Pelargopsis (9 sp.), the whole Oriental region, and extending to Celebes and Timor in the Austro-Malayan subregion; Ceya (11 sp.), the Oriental region and Austro-Malayan subregion, but absent from Celebes, and only one species in continental India and Ceylon; Ceycopsis (1 sp.), Celebes; Myioceyx (2 sp.), West Africa ; Ipsidina (4 sp.), Ethiopian region ; Syma (2 sp.), Papua and North Australia; Halcyon (36 sp.), Australian, Oriental, and Ethiopian regions, and the southern part of the Palæarctic; Dacelo (6 sp.), Australia and New Guinea ; Todir*hamphus (3 sp.), Eastern Pacific Islands only; Monachalcyon (1 sp.), Celebes; Caridonax (1 sp.), Lombok and Flores; Corcineutes (2 sp.), Siam to Borneo and Java ; Tanysiptera (14 sp.), Moluccas New Guinea, and North Australia (Plate X. Vol. I. p. 414); Cittura (2 sp.), Celebes group; Melidora (1 sp.), New Guinea.

FAMILY 68.—BUCEROTIDIÆ. (12 Genera, 50 Species.)

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The Hornbills form an isolated group of generally large-sized birds, whose huge bills form their most prominent feature. They are popularly associated with the American Toucans, but have no close relationship to them, and are now generally

1958

considered to show most resemblance, though still a very distant one, to the kingfishers. They are abundant in the Ethiopian and Oriental regions, and extend eastward to the Solomon Islands. Their classification is very unsettled, for though they have been divided into more than twenty genera they have not yet been carefully studied. The following grouping of the genera—referring to the numbers in the Hand List—must therefore be considered as only provisional :

(1957 1963) Buceros (6 sp.), all Indo-Malaya, Arakan, Nepal and the Neilgherries (Plate IX. Vol. I. p. 339); (1969 - 1961) Hydrocissa (7 sp.), India and Ceylon to Malaya and Celebes ; (1962) Berenicornis (2 sp.), Sumatra and West Africa; (1964) Calao (3 sp.), Tennaserim, Malaya, Moluccas to the Solomon Islands ; (1385) Aceros (1 sp.), South-east Himalayas ; (1966 1967) Cranorrhinus (3 sp.), Malacca, Sumatra, Borneo, Philippines, Celebes; (1968) Penelopides (1 sp.), Celebes; (1969 – 1971) Tockus (15 sp.), Tropical and South Africa ; (1972) Rhinoplax (1 sp.), Sumatra and Borneo; (1973 – 1975) Bycanistes (6 sp.), West Africa with East and South Africa ; (1976 1977) Meniceros (3 sp.), India and Ceylon to Tenasserim; (1978) Bucorvus (2 sp.), Tropical and South Africa.

FAMILY 69.—UPUPIDÆ. (1 Genus, 6 Species.)

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The Hoopoes form a small and isolated group of semi-terrestrial insectivorous birds, whose nearest affinities are with the Hornbills. They are most characteristic of the Ethiopian region, but extend into the South of Europe and into all the continental divisions of the Oriental region, as well as to Ceylon, and northwards to Pekin and Mongolia.

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