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It has already been shown (Vol. I. pp. 209-213 and Vol. II. pp. 44-48) that the peculiarities of distribution of the various groups of insects depend very much on their habits and general economy. Their antiquity is so vast, and their more important modifications of structure have probably occurred so slowly, that modes of dispersal depending on such a combination of favourable conditions as to be of excessive rarity, may yet have had time to produce large cumulative effects. Their small specific gravity and their habits of flight render them liable to dispersal by winds to an extent unknown in other classes of animals; and thus, what are usually very effectual barriers have been overstepped, and sometimes almost obliterated, in the case of insects. A careful examination will, however, almost always show traces of an ancient fauna, agreeing in character with other classes of animals, intermixed with the more prominent and often more numerous forms whose presence is due to this unusual facility of dispersal.
The effectual migration of insects is, perhaps more than in any other class of animals, limited by organic and physical conditions. The vegetation, the soil, the temperature, and the supply of moisture, must all be suited to their habits and economy; while they require an immunity from enemies of various kinds, which immigrants to a new country seldom obtain. Few organisms have, in so many complex ways, become adapted to their special environment, as have insects. They are in each country more or less adapted to the plants which belong to it; while their colours, their habits, and the very nature of the juices of their system, are all modified so as to protect them from the special dangers which surround them in their native land. It follows, that while no animals are so well adapted to show us the various modes by which dispersal may be effected, none can so effectually teach us the true nature and vast influence of the organic barrier in limiting dispersal.
It is probable that insects have at one time or another taken advantage of every line of migration by which any terrestrial organisms have spread over the earth, but owing to their small size and rapid multiplication, they have made use of some which are exclusively their own. Such are the passage along mountain ranges from the Arctic to the Antarctic regions, and the dispersal of certain types over all temperate lands. It will perhaps be found that insects have spread over the land surface in directions dependent on our surface zones—forests, pastures, and deserts;-and a study of these, with a due consideration of the fact that narrow seas are scarcely a barrier to most of the groups, may assist us to understand many of the details of insect-distribution.
The distribution of land-shells agrees, in some features, with that of insects, while in others the two are strongly contrasted. In both we see the effects of great antiquity, with some special means of dispersal; but while in insects the general powers of motion, both voluntary and involuntary, are at a maximum, in land-molluscs they are almost at a minimum. Although to some extent dependent on vegetation and climate, the latter are more dependent on inorganic conditions, and also to a large extent on the general organic environment. The result of these various causes, acting through countless ages, has been to spread the main types of structure with considerable uniformity over the globe ; while generic and sub-generic forms are often wonderfully localized. Land-shells, even more than insects, seem, at first sight, to require regions of their own; but we have already pointed out the disadvantages of such a method of study. It will be far more instructive to refer them to those regions and sub-regions which are found to accord best with the distribution of the higher animals, and to consider the various anomalies they present as so many problems, to be solved by a careful study of their habits and economy, and especially by a search after the hidden causes which have enabled them to spread so widely over land and ocean. The lines of migration which land-shells have followed, can WOL. II.-36
hardly be determined with any definiteness. On continents they seem to spread steadily, but slowly, in every direction, checked probably by organic and physical conditions rather than by the barriers which limit the higher groups. Over the ocean they are also slowly dispersed, by some means which act perhaps at very long intervals, but which, within the period of the duration of genera and families, are tolerably effective. It thus happens that, although the powers of dispersal of land-shells and insects are so very unequal, the resulting geographical distribution is almost the opposite of what might have been expected,—the former being, on the whole, less distinctly localized than the latter.
The preceding remarks are all I now venture to offer, on the distinguishing features of the various groups of land-animals as regards their distribution and migrations. They are at best but indications of the various lines of research opened up to us by the study of animals from the geographical point of view, and by looking upon their range in space and time as an important portion of the earth's history. Much work has yet to be done before the materials will exist for a complete treatment of the subject in all its branches; and it is the author's hope that his volumes may lead to a more systematic collection and arrangement of the necessary facts. At present all public museums and private collections are arranged zoologically. All treatises, monographs, and catalogues, also follow, more or less completely, the zoological arrangement; and the greatest difficulty the student of geographical distribution has to contend against, is the total absence of geographical collections, and the almost total want of complete and comparable local catalogues. Till every wellmarked district, every archipelago, and every important island, has all its known species of the more important groups of animals catalogued on a uniform plan, and with a uniform nomenclature, a thoroughly satisfactory account of the Geographical Distribution of Animals will not be possible. But more than this is wanted. Many of the most curious relations between animal forms and their habitats, are entirely unnoticed, owing to the productions of the same locality never being associated in our museums and collections. A few such relations have been brought to light by modern scientific travellers, but many more remain to be discovered; and there is probably no fresher and more productive field still unexplored in Natural History. Most of these curious and suggestive relations are to be found in the productions of islands, as compared with each other, or with the continents of which they form appendages; but these can never be properly studied, or even discovered, unless they are visibly grouped together. When the birds, the more conspicuous families of insects, and the land-shells of islands, are kept together so as to be readily compared with similar associations from the adjacent continents or other islands, it is believed that in almost every case there will be found to be peculiarities of form or colour running through widely different groups, and strictly indicative of local or geographical influences. Some of these coincident variations have been alluded to in various parts of this work, but they have never been systematically investigated. They constitute an unworked mine of wealth for the enterprising explorer; and they may not improbably lead to the discovery of some of the hidden laws (supplementary to Natural Selection), which seem to be required, in order to account for many of the external characteristics of animals.
In concluding his task, the author ventures to suggest that naturalists who are disposed to turn aside from the beaten track of research, may find in the line of study here suggested a new and interesting pursuit, not inferior in attractions to the lofty heights of transcendental anatomy, or the bewildering mazes of modern classification. And it is a study which will surely lead them to an increased appreciation of the beauty and the barmony of nature, and to a fuller comprehension of the complex relations and mutual interdependence, which link together every animal and vegetable form, with the ever-changing earth which supports them, into one grand organic whole.