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It is a fact within the experience of most persons, that the various species of animals are not uniformly dispersed over the surface of the country. If we have a tolerable acquaintance with any district, be it a parish, a county, or a larger extent of territory, we soon become aware that each well-marked portion of it has some peculiarities in its animal productions. If we want to find certain birds or certain insects, we have not only to choose the right season but to go to the right place.
If we travel beyond our district in various directions we shall almost certainly meet with something new to us; some species which we were accustomed to see almost daily will disappear, others which we have never seen before will make their appearance. If we go very far, so as to be able to measure our journey by degrees of latitude and longitude and to perceive important changes of climate and vegetation, the differences in the forms of animal life will become greater ; till at length we shall come to a country where almost everything will be new, all the familiar creatures of our own district being replaced by others more or less differing from them.
If we have been observant during our several journeys, and have combined and compared the facts we have collected, it will become apparent that the change we have witnessed has been of two distinct kinds. In our own and immediately surrounding districts, particular species appeared and disappeared because
the soil, the aspect, or the vegetation, was adapted to them or the reverse. The marshes, the heaths, the woods and forests, the chalky downs, the rocky mountains, had each their peculiar inhabitants, which reappeared again and again as we came to tracts of country suitable for them. But as we got further away we began to find that localities very similar to those we had left behind were inhabited by a somewhat different set of species; and this diflerence increased with distance, notwithstanding that almost identical external conditions might be often met with. The first class of changes is that of stations; the second that of habitats. The one is a local, the other a geographical phenomenon. The whole area over which a particular animal is found may consist of any number of stations, but rarely of more than one habitat. Stations, however, are often so extensive as to include the entire range of many species. Such are the great seas and oceans, the Siberian or the Amazonian forests, the North African deserts, the Andean or the Himalayan highlands.
There is yet another difference in the nature of the change we have been considering. The new animals which we meet with as we travel in any direction from our starting point, are some of them very much like those we have left behind us, and can be at once referred to familiar types; while others are altogether unlike anything we have seen at home. When we reach the Alps we find another kind of squirrel, in Southern Italy a distinct mole, in Southern Europe fresh warblers and unfamiliar buntings. We meet also with totally new forms; as the glutton and the snowy owl in Northern, the genet and the hoopoe in Southern, and the saiga antelope and collared pratincole in Eastern Europe. The first series are examples of what are termed representative species, the second of distinct groups or types of animals. The one represents a comparatively recent modification, and an origin in or near the locality where it occurs; the other is a result of very ancient changes both organic and inorganic, and is connected with some of the most curious and difficult of the problems we shall have to discuss.
Having thus defined our subject, let us glance at the opinions that have generally prevailed as to the nature and causes of the phenomena presented by the geographical distribution of animals.
It was long thought, and is still a popular notion, that the manner in which the various kinds of animals are dispersed over the globe is almost wholly due to diversities of climate and of vegetation. There is indeed much to favour this belief. The arctic regions are strongly characterised by their white bears and foxes, their reindeer, ermine, and walruses, their white ptarmigan, owls, and falcons; the temperate zone has its foxes and wolves, its rabbits, sheep, beavers, and marmots, its sparrows and its song birds; while tropical regions alone produce apes and elephants, parrots and peacocks, and a thousand strange quadrupeds and brilliant birds which are found nowhere in the cooler regions. So the camel, the gazelle and the ostrich live in the desert; the bison on the prairie; the tapir, the deer, and the jaguar in forests. Mountains and marshes, plains and rocky precipices, have each their animal inhabitants; and it might well be thought, in the absence of accurate inquiry, that these and other differences would sufficiently explain why most of the regions and countries into which the earth is popularly divided should have certain animals peculiar to them and should want others which are elsewhere abundant.
A more detailed and accurate knowledge of the productions of different portions of the earth soon showed that this explanation was quite insufficient; for it was found that countries exceedingly similar in climate and all physical features may yet have very distinct animal populations. The equatorial parts of Africa and South America, for example, are very similar in climate and are both covered with luxuriant forests, yet their animal life is widely different; elephants, apes, leopards, guinea-fowls and touracos in the one, are replaced by tapirs, prehensiletailed monkeys, jaguars, curassows and toucans in the other. Again, parts of South Africa and Australia are wonderfully similar in their soil and climate; yet one has lions, antelopes, zebras and giraffes ; the other only kangaroos, wombats, phalan
gers and mice. In like manner parts of North America and Europe are very similar in all essentials of soil climate and vegetation, yet the former has racoons, opossums, and hummingbirds; while the latter possesses moles, hedgehogs and true flycatchers. Equally striking are the facts presented by the distribution of many large and important groups of animals. Marsupials (opossums, phalangers &c.) are found from temperate Van Diemen's land to the tropical islands of New Guinea and Celebes, and in America from Chili to Virginia. No crows exist in South America, while they inhabit every other part of the world, not excepting Australia. Antelopes are found only in Africa and Asia ; the sloths only in South America ; the true lemurs are confined to Madagascar, and the birds-of-paradise to New Guinea
If we examine more closely the distribution of animals in any extensive region, we find that different, though closely allied species, are often found on the opposite sides of any considerable barrier to their migration. Thus, on the two sides of the Andes and Rocky Mountains in America, almost all the mammalia, birds, and insects are of distinct species. To a less extent, the Alps and Pyrenees form a similar barrier, and even great rivers and river plains, as those of the Amazon and Ganges, separate more or less distinct groups of animals. Arms of the sea are still more effective, if they are permanent; a circumstance in some measure indicated by their depth. Thus islands far away from land almost always have very peculiar animals found nowhere else; as is strikingly the case in Madagascar and New Zealand, and to a less degree in the West India islands. But shallow straits, like the English Channel or the Straits of Malacca, are not found to have the same effect, the animals being nearly or quite identical on their opposite shores. A change of climate or a change of vegetation may form an equally effective barrier to migration. Many tropical and polar animals are pretty accurately limited by certain isothermal lines; and the limits of the great forests in most parts of the world strictly determine the ranges of many species.
Naturalists have now arrived at the conclusion, that by some