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mammalia of North America are not very numerous; and they can be for the most part divided into two groups, the one allied to the Palearctic, and the other to the Neotropical fauna. The bears, the wolves, the cats, the bison, sheep and antelope, the hares, the marmots, and the pikas, resemble Palearctic forms; while the racoons, skunks, opossum, and vesper-mice are now

more peculiarly Neotropical. There are also many genera which are altogether peculiar and characteristic of the region, as the prong-horn antelope (Antilocapra), the jumping-mouse (Jaculus), five genera of pouched rats (Saccomyidae), the prairie dogs (Cynomys), the tree porcupines (Erethizon), and some others.

Birds present the same mixture of the two types; but the wild turkeys (Meleagris), the passenger pigeon (Ectopistes), the crested quails (Lophortyx, &c.), the ruffed grouse (Cupidonia), and some other groups of less importance, are peculiar; while the family of the wood warblers (Mniotiltide) is so largely developed that it may claim to be more characteristic of North than of South America.

Reptiles and Amphibia present a number of peculiar types; while no less than five peculiar families of freshwater fishes would alone serve to mark out this as distinct from every other part of the world.

Considering the evident affinity between the Nearctic and Palearctic regions, there are here some curious deficiencies of groups which are common and widelyspread in the latter. Thus hedgehogs, wild horses and asses, swine, true oxen, goats, dormice, and true mice are absent; while sheep and antelopes are only represented by solitary species in the Rocky Mountains.

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Among birds, too, we have such striking deficiencies as the extensive families of flycatchers, starlings, and pheasants.

Turning now to the Neotropical region, comprising all South America and the tropical parts of the northern continent, we find that the Old World types have still further diminished, while a number of new and altogether peculiar forms have taken their place. Insectivora have wholly disappeared with the exception of one anomalous form in the greater Antilles; bears are represented by one Chilian species; swine are replaced by peccaries; the great Bovine family are entirely unknown; the camel tribe are confined to the Southern Andes and the south temperate plains; deer are not numerous; and all the varied Ungulata of the Old World are represented only by a few species of tapirs. These great gaps are, however, to some extent filled up by a variety of interesting and peculiar types. Two families of monkeys (Cebidae and Hapalide) differ in many points of structure from all the Quadrumana of the eastern hemisphere. There is a peculiar family of bats the vampyres; many peculiar weasels and Procyonidae; a host of peculiar rodents, comprising five distinct families, among which are the largest living forms of the order; and a great number of Edentata, comprising the families of the sloths, armadillos, and ant-eaters; and lastly, a considerable number of the marsupial family of opossums. As compared with the Old World, we find here a great abundance and variety of the lower types, with a corresponding scarcity of such higher forms as characterise the tropics of Africa and Asia.

In birds we meet with corresponding phenomena. The most abundant and characteristic families of the Old World tropics are replaced here by a series of families of a lower grade of organisation, among which are such remarkable groups as the chatterers (Cotingida), the manakins (Pipridae), the ant-thrushes (Formicariidae), the toucans (Rhamphastida), the motmots (Momotidae), and the humming-birds (Trochilidae), the last perhaps the most remarkable and beautiful of all developments of the bird-type. Parrots are numerous, but these, too, are mostly of peculiar families; while pheasants and grouse are replaced by curassows and tinamous, and there are an unusual number of remarkable and isolated forms of waders.

Reptiles, amphibia, fresh-water fishes, insects, and land-shells, are all equally peculiar and abundant; so that South America presents, on the whole, an assemblage of curious and beautiful natural objects, unsurpassedperhaps even unequalled-in any other part of the globe.

Past History of the American Continents.—We will now proceed to examine what is known of the past history of the two American continents, and endeavour to determine what have been their former relations to each other and to the Old World, and how their existing zoological and geographical features have been brought about. And first let us see what knowledge we possess of the past relations of North America with the Eastern continents.

If we go back to that recent period termed the PostPliocene-corresponding nearly to the Post-Glacial period and to that of pre-historic man in Europe-we

find at once a nearer approximation than now exists. between the Nearctic and Palearctic faunas. North America then possessed several large cats, six distinct species of the horse family, a camel, two bisons, and four species of elephants and mastodons. A little earlier, in the Pliocene period (although fossil remains of this age are scanty), we have in addition the genus Rhinoceros several distinct camels, some new forms of ruminants and an Old-World form of porcupine. Further back, in the Miocene period, we find a Lemuroid animal, numerous insectivora, a host of carnivora, chiefly feline and canine, a variety of equine and tapirine forms, rhinoceroses, camels, deer, and an extensive extinct family-the Oreodontida-allied to deer, camels, and swine. There are, however, no elephants. In the still earlier Eocene period most of the animals were peculiar, and unlike anything now living, but some were identical with European types of the same age, as Lophotherium and the family Anchitherida.

These facts compel us to believe that at distinct epochs during the Tertiary period the interchange of large mammalia between North America and the Old World has been far more easy than it is now. In the Post-Pliocene period, for example, the horses, elephants, and camels of North America and Europe were so closely allied that their common ancestors must have passed from one continent to the other, just as we feel assured that the common ancestors of the American and European bison, elk, and beaver, must have so migrated. We have further evidence in the curious fact that certain groups appear to come into existence in the one continent much later than in the other. Thus cats, deer, masto

dons, true horses, porcupines, and beavers, existed in Europe long before they appeared in America; and as the theory of evolution does not admit the independent development of the same group in two disconnected regions to be possible, we are forced to conclude that these animals have migrated from one continent to the other. Camels, and perhaps ancestral horses, on the other hand, were more abundant and more ancient in America, and may have migrated thence into Northern Asia

There are two probable routes for such migrations. From Norway to Greenland by way of Iceland and across Baffin Bay to Arctic America, there is everywhere a comparatively shallow sea, and it is not improbable that during the Miocene period, or subsequently, a land communication may have existed here. On the other side of the continent, at Behring Straits, the probability is greater. For here we have a considerable extent of far shallower sea, which a very slight elevation would convert into a broad isthmus connecting North America and North-East Asia. It is true that elephants, horses, deer, and camels would, under existing climatal conditions, hardly range as far north as Greenland and Alaska; but we must remember that most mysterious yet indisputable fact of the luxuriant vegetation, including even magnolias and other large-leaved evergreens, which flourished in these latitudes during the Miocene period; so that we have all the conditions of favourable climate and abundant food, which would render such interchange of the animals of the two continents not only possible, but inevitable, whenever a land communication was effected; and there is reason

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