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enough sense to get on to the lower sides of the leaves if it begins to rain, and from time to time changes its skin. This is its whole existence, which certainly does not lead us to expect a display of any, even the most limited, intellectual power. When, however, the time comes for the larva of this moth to become a chrysalis, it spins for itself a double cocoon, fortified with bristles that point outwards, so that it can be opened easily from within, though it is sufficiently impenetrable from without. If this contrivance were the result of conscious reflection, we should have to suppose some such reasoning process as the following to take place in the mind of the caterpillar :—“ I am about to become a chrysalis, and, motionless as I must be, shall be exposed to many different kinds of attack. I must therefore weave myself a web. But when I am a moth I shall not be able, as some moths are, to find my way out of it by chemical or mechanical means; therefore I must leave a way open for myself. In order, however, that my enemies may not take advantage of this, I will close it with elastic bristles, which I can easily push asunder from within, but which, upon the principle of the arch, will resist all pressure from without." Surely this is asking rather too much from a

poor caterpillar; yet the whole of the foregoing must be thought out if a correct result is to be arrived at.

This theoretical separation of instinct from conscious intelligence can be easily misrepresented by opponents of my theory, as though a separation in practice also would be necessitated in consequence. This is by no means my intention. On the contrary, I have already insisted at some length that both the two kinds of mental activity may co-exist in all manner of different proportions, so that there may be every degree of combination, from pure instinct to pure deliberation. We shall see, however, in a later chapter, that even in the highest and most abstract activity of human consciousness there are forces at work that are of the highest importance, and are essentially of the same kind as instinct.

On the other hand, the most marvellous displays of instinct are to be found not only in plants, but also in those lowest organisms of the simplest bodily form which are partly unicellular, and in respect of conscious intelligence stand far below the higher plants—to which, indeed, any kind of deliberative faculty is commonly denied. Even in the case of those minute microscopic organisms that baffle our

attempts to classify them either as animals or vegetables, we are still compelled to admire an instinctive, purposive behaviour, which goes far beyond a mere reflex response to a stimulus from without; all doubt, therefore, concerning the actual existence of an instinct must be at an end, and the attempt to deduce it as a consequence of conscious deliberation be given up as hopeless. I will here adduce an instance as extraordinary as any we yet know of, showing, as it does, that many different purposes, which in the case of the higher animals require a complicated system of organs of motion, can be attained with incredibly simple means.

Arcella vulgaris is a minute morsel of protoplasm, which lives in a concave - convex, brown, finely reticulated shell, through a circular opening in the concave side of which it can project itself by throwing out pseudopodia. If we look through the microscope at a drop of water containing living arcella, we may happen to see one of them lying on its back at the bottom of the drop, and making fruitless efforts for two or three minutes to lay hold of some fixed point by means of a pseudopodium. After this there will appear suddenly from two to five, but sometimes more, dark points in the protoplasm at a small distance from the

circumference, and, as a rule, at regular distances from one another. These rapidly develop themselves into well-defined spherical air vesicles, and come presently to fill a considerable part of the hollow of the shell, thereby driving part of the protoplasm outside it. After from five to twenty minutes, the specific gravity of the arcella is so much lessened that it is lifted by the water with its pseudopodia, and brought up against the upper surface of the water-drop, on which it is able to travel. In from five to ten minutes the vesicles will now disappear, the last small point vanishing with a jerk. If, however, the creature has been accidentally turned over during its journey, and reaches the top of the water-drop with its back uppermost, the vesicles will continue growing only on one side, while they diminish on the other; by this means the shell is brought first into an oblique and then into a vertical position, until one of the pseudopodia obtains a footing and the whole turns over. From the moment the animal has obtained foothold, the bladders become immediately smaller, and after they have disappeared the experiment may be repeated at pleasure.

The positions of the protoplasm which the vesicles fashion change continually; only the

grainless protoplasm of the pseudopodia develops no air. After long and fruitless efforts a manifest fatigue sets in; the animal gives up the attempt for a time, and resumes it after an interval of repose.

Engelmann, the discoverer of these phenomena, says (Pflüger's Archiv für Physologie, Bd. II.): The changes in volume in all the vesicles of the same animal are for the most part synchronous, effected in the same manner, and of like size. There are, however, not a few exceptions; it often happens that some of them increase or diminish in volume much faster than others; sometimes one may increase while another diminishes; all the changes, however, are throughout unquestionably intentional. The object of the air-vesicles is to bring the animal into such a position that it can take fast hold of something with its pseudopodia. When this has been obtained, the air disappears without our being able to discover any other reason for its disappearance than the fact that it is no longer needed. . . . If we bear these circumstances in mind, we can almost always tell whether an arcella will develop air-vesicles or no; and if it has already developed them, we can tell whether they will increase or diminish. ... The arcella, in fact, in this power of alter

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