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of organic beings show, that both these plans have obtained in creation. Sometimes the series of affinities can be well represented for a space by a direct progression from species to species or from group to group, but it is generally found impossible so to continue. There constantly occur two or more modifications of an organ or modifications of two distinct organs, leading us on to two distinct series of species, which at length differ so much from each other as to form distinct genera or families. These are the parallel series or representative groups of naturalists, and they often occur in different countries, or are found fossil in different formations. They are said to have an analogy to each other when they are so far removed from their common antitype as to differ in many important points of structure, while they still preserve a family resemblance. We thus see how difficult it is to determine in every case whether a given relation is an analogy or an affinity, for it is evident that as we go back along the parallel or divergent series, towards the common antitype, the analogy which existed between the two groups becomes an affinity. We are also made aware of the difficulty of arriving at a true classification, even in a small and perfect group ;—in the actual state of nature it is almost impossible, the species being so numerous and the modifications of form and structure so varied, arising probably from the immense number of species which have served as antitypes for the existing species, and thus produced a complicated branching of the lines of affinity, as intricate as the twigs of a gnarled oak or the vascular system of the human body. Again, if we consider that we have only fragments of this vast system, the stem and main branches being represented by extinct species of which we have no knowledge, while a vast mass of limbs and boughs and minute twigs and scattered leaves is what we have to place in order, and determine the true position each originally occupied with regard to the others, the whole difficulty of the true Natural System of classification becomes apparent to us. We shall thus find ourselves obliged to reject all those systems of classification which arrange species or groups in circles, as well as those which fix a definite number for the divisions of each group. The latter class have been very generally rejected by naturalists, as contrary to nature, notwithstanding the ability with which they have been advocated ; but the circular system of affinities seems to have obtained a deeper hold, many eminent naturalists having to some extent adopted it. We have, however, never been able to find a case in which the circle has been closed by a direct and close affinity. In most cases a palpable analogy has been substituted, in others the affinity is very obscure or altogether doubtful. The complicated branching of the lines of affinities in extensive groups must also afford great facilities for giving a show of probability to any such purely artificial arrangements. Their death-blow was given by the admirable paper of the lamented Mr. Strickland, published in the “Annals of Natural History,” in which he so clearly showed the
true synthetical method of discovering the Natural System.
Geographical Distribution of Organisms.
If we now consider the geographical distribution of animals and plants upon the earth, we shall find all the facts beautifully in accordance with, and readily explained by, the present hypothesis. A country having species, genera, and whole families peculiar to it, will be the necessary result of its having been isolated for a long period, sufficient for many series of species to have been created on the type of pre-existing ones, which, as well as many of the earlierformed species, have become extinct, and thus made the groups appear isolated. If in any case the antitype had an extensive range, two or more groups of species might have been formed, each varying from it in a different manner, and thus producing several representative or analogous groups. The Sylviadae of Europe and the Sylvicolidae of North America, the Heliconidae of South America and the Euploeas of the East, the group of Trogons inhabiting Asia, and that peculiar to South America, are examples that may be accounted for in this manner.
Such phaenomena as are exhibited by the Galapagos Islands, which contain little groups of plants and animals peculiar to themselves, but most nearly allied to those of South America, have not hitherto received any, even a conjectural explanation. The Galapagos are a volcanic group of high antiquity, and have probably never been more closely connected with the continent than they are at present. They must have been first peopled, like other newly-formed islands, by the action of winds and currents, and at a period sufficiently remote to have had the original species die out, and the modified prototypes only remain. In the same way we can account for the separate islands having each their peculiar species, either on the supposition that the same original emigration peopled the whole of the islands with the same species from which differently modified prototypes were created, or that the islands were successively peopled from each other, but that new species have been created in each on the plan of the pre-existing ones. St. Helena is a similar case of a very ancient island having obtained an entirely peculiar, though limited, flora. On the other hand, no example is known of an island which can be proved geologically to be of very recent origin (late in the Tertiary, for instance), and yet possesses generic or family groups, or even many species peculiar to itself. When a range of mountains has attained a great elevation, and has so remained during a long geological period, the species of the two sides at and near their bases will be often very different, representative species of some genera occurring, and even whole genera being peculiar to one side only, as is remarkably seen in the case of the Andes and Rocky Mountains. A similar phaenomenon occurs when an island has been separated from a continent at a very early period. The shallow sea between the Peninsula of Malacca, Java, Sumatra and Borneo was probably a continent or large island at an early epoch, and may have become submerged as the volcanic ranges of Java and Sumatra were elevated. The organic results we see in the very considerable number of species of animals common to some or all of these countries, while at the same time a number of closely allied representative species exist peculiar to each, showing that a considerable period has elapsed since their separation. The facts of geographical distribution and of geology may thus mutually explain each other in doubtful cases, should the principles here advocated be clearly established. In all those cases in which an island has been separated from a continent, or raised by volcanic or coralline action from the sea, or in which a mountain-chain has been elevated in a recent geological epoch, the phaenomena of peculiar groups or even of single representative species will not exist. Our own island is an example of this, its separation from the continent being geologically very recent, and we have consequently scarcely a species which is peculiar to it; while the Alpine range, one of the most